While seeking out the Clairvoyant the team boards a train that appears to be heading toward doom. In an attempt to save Skye Coulson enlists the help of Ward's former S. John Garrett Bill Paxton ; shocking truths are uncovered. Garrett and Triplett help search for the Clairvoyant but face the wrath of Deathlok.
Coulson and the team are trapped with a traitor in their midst without anyone they can trust. With Colonel Glenn Talbot now on their trail Coulson and his team seek refuge in the last place anyone would look where they begin to uncover S.
With their world turned upside down Coulson races to save the life of his one true love as the mystery of The Cellist which began in Marvel's The Avengers. In the last episode before the epic season finale Ward's betrayal and HYDRA's shocking secrets are revealed as Coulson's team goes undercover on a mission that leaves no one unscathed.
On the season premiere Coulson and his team are now wanted fugitives with limited resources--but that's not stopping them from keeping the world safe from powerful and unseen threats everywhere.
However with new members they hardly know will S. Hunted and running out of solutions Coulson puts his team right in the line of fire in the hopes of saving them all. But with General Talbot Hydra and Creel all on the attack can they possibly survive?
Meanwhile a mysterious doctor harbors some dangerous secrets that could destroy one of the core team members. Coulson and his team race against Hydra to get to Donnie Gill--a dangerous gifted with the ability to freeze objects--while Simmons is caught in the crossfire.
While on a mission to learn more about the secrets of the mysterious writing Coulson finds himself attacked by the only person he can trust: Agent Melinda May. Meanwhile the rest of the team is trapped in an explosive situation and it's up to Fitz to save them.
Meanwhile Skye's father forces Raina to reunite him with his daughter at any cost. The world turns against S. The team discovers murder victims with the alien glyphs on them that Coulson's been etching. All of Coulson's secrets and lies come to a head. Coulson and team find themselves in an epic face-off against Hydra to uncover an ancient secret while Ward kidnaps his brother Senator Christian Ward for a violent trip down memory lane. SHIELD discovers the ancient city before Hydra but uncovering the secrets may require one of Coulson's team to make the ultimate sacrifice.
Meanwhile May and Skye race to get to Raina before Whitehall takes her. Coulson and Whitehall's forces meet in an explosive confrontation that dramatically alters everyone's fates. Meanwhile Skye discovers shocking secrets about her past on the winter finale. After discovering an alien city with ties to his resurrection Coulson and his team destroyed it before the forces of Hydra could claim its secrets eliminating the villainous Whitehall in the process.
But new threats to the world have arisen including Skye's father Cal who now seeks retribution against Coulson for stealing his revenge against Whitehall.
Coulson and team come to the aid of a confused Lady Sif who has lost her memory after fighting with a mysterious warrior whose true mission will send shockwaves through S. Meanwhile Bobbi and Mac continue to hide their true mission from everyone close to them.
Cal seeks revenge on Coulson by assembling a team of super villains to destroy S. Meanwhile May calls on renowned Dr. Andrew Garner her charismatic ex-husband with a crisis that threatens to tear the team apart.
Coulson and his team are still reeling from a shocking revelation that leaves the team fractured as they must decide what to do with one of their own. Meanwhile Bobbi and Mack reveal their true allegiance to Hunter while elsewhere Ward and Agent 33 embark on a personal mission. War comes to Coulson's doorstep in a way he never anticipated as shocking revelations are brought to light and Skye struggles to control her new abilities but will soon make a decision that irrevocably changes her life.
As Robert Gonzales makes his move Coulson must do whatever it takes to protect the future of S. Meanwhile Skye's journey to control her powers takes a surprising turn when she meets the enigmatic Inhuman named Lincoln. The story of Melinda May's transformation into The Cavalry is revealed as she finds herself at a crossroads in the war between Coulson and Gonzales. As Skye learns more about her true heritage from Lincoln the truth about the Inhumans will send her in a new direction.
A dangerous new alliance is formed as Coulson and Hunter turn to the last person anyone would expect--Grant Ward! As the war between S. Gonzales and Coulson must find a way to put their differences aside and work together against Hydra even if it means teaming up with someone they don't trust.
Skye is torn between her loyalty to S. In part one of the season finale S. Coulson and his team will be forced to make shocking sacrifices that will leave their relationships and their world changed forever. In part two of the season finale S. When Coulson and the team discover a new Inhuman S. And still reeling from Simmons' dramatic disappearance Fitz goes to extreme lengths to try to learn how to get her back.
Fitz and the team enlist the aid of an Asgardian to unlock the secrets of the ancient monolith that swallowed Simmons and Agent May is at a crossroads in her personal and professional life. Daisy Mack and Coulson race to protect Lincoln as Rosalind's team hunts down the Inhuman in their quest for powered people. Coulson makes a desperate move to save his team as multiple enemies close in. Meanwhile, a doctor harbors secrets that could threaten one of the agents.
A team member is caught in the middle as Coulson and the agents try to reach a man with dangerous freezing powers before Hydra does.
Coulson seeks to unlock the mystery of his alien writing and is attacked by a trusted ally. Meanwhile, with the team trapped, Fitz might be their only hope. The team faces off against Hydra security chief Bobbi Morse. Ward's vendetta and plan for revenge against S. D battles Hydra head-to-head, which sees Coulson and Fitz take the ultimate risk while Daisy and the Inhumans try to keep Hydra at bay.
Coulson is more determined than ever to get to Gideon Malick and put to an end to Hydra once and for all in the aftermath of his trip to Maveth. Meanwhile, Daisy and the team encounter more Inhumans who have powers like they've never seen before, but will they be friends or enemies of S.
Coulson and the team investigate an international conference on the fate of the Inhumans. Back on the bus, Fitzsimmons may have found a way to stop the Inhuman gene from spreading. After tracking Malick to Russia, Bobbi and Hunter become involved in an assassination plot; the team is changed forever as they race to save lives in the balance.
Agent Mack and his brother step in when a radical group called the Watchdogs plan to eliminate the Inhumans; Simmons finds a powerful chemical compound. When Daisy gets a horrifying glimpse of the future, S. The team hunts for information on a new enemy; Malick's secret from his past threatens to destroy his Hydra legacy. Agent Daisy Johnson must call upon the Secret Warriors for an inaugural mission that will leave no member unscathed, and S.
The S. But there is a sliver of hope as Agents Fitz and Simmons follow a lead that may be able to stop the maniacal Inhuman once and for all.
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